Cellular growth signals are often transmitted through a process called phosphorylation, in which an enzyme called tyrosine kinase attaches a phosphate tag to another protein. This phosphate tag is derived from a molecule called ATP.2
EGFR is a protein in the receptor tyrosine kinase family. This protein consists of a receptor domain outside the cell connected to a tyrosine kinase domain inside the cell.3
The kinase domain of EGFR is inactive by itself, but when two kinase domains come together to form a dimer, the kinase is activated. Active EGFR dimer then phosphorylates itself to transmit growth signals.4
In normal conditions, EGFR does not associate with each other and remains monomeric and inactive.4
On the other hand, in the presence of a growth factor called EGF3, the structure of the receptor domain of EGFR changes in a way that facilitates dimerization of EGFR.5,6
This brings two kinase domains of EGFR together and activates them to transmit growth signals through phosphorylation.4 Of note, because the dimerization of EGFR requires the binding of growth factors, growth signaling transmitted by EGFR is controllable; when cell growth is no longer necessary, growth factor production ceases, which returns EGFR to its inactive monomeric form.4
In some types of cancer, a mutation in EGFR results in the activation of EGFR in the absence of growth factors. Some mutations cause EGFR to dimerize without growth factors, while other mutations cause the EGFR kinase to be active in its monomeric form.7 In either case, the kinases of EGFR are constitutively active.
Gefitinib is a type of drug called a kinase inhibitor. It blocks the kinase domain of EGFR from accessing the ATP required for phosphorylation.8 This turns off the growth signal, and as a result, cancer growth is halted.